Prof. Shihui Han and the Culture and Social Cognitive Neuroscience lab study cultural and genetic influences on psychological and neural mechanisms underlying social cognition and social behavior by integrating social/cultural psychology and brain imaging such as fMRI/EEG/MEG. Their research aims to clarify how the brain represents self-related information (self-referential processing), how the brain understands and shares others’ emotion (empathy), and how the brain processes death-related information and the related effects on other social cognitive/affective processes.

List of Publications


Han, X., Gelfand, M. J., Wu, B., Zhang, T., Li, W., Gao, T., Pang, C.,Wu, T., Zhou, Y., Zhou, S., Wu, X., Han, S. (2020). A neurobiological association of revenge propensity during intergroup conflict. eLife, in press.

Luo, S., Zhu, Y., Fan, L., Gao, D., Han, S. (2020). Resting-state brain network properties mediate the association between the oxytocin receptor gene and interdependence. Social Neuroscience, in press.

Gao, T., Zhou, Y., Li, W., Pfabigan, D. M., Han, S. (2019). Neural mechanisms of reinforcement learning under mortality threat. Social Neuroscience, in press.

Vu, T., van der Meulen, A., Heslenfeld, D., Woodcock, K., Han, S., & Krabbendam, L. (2020). Neural responses to affective stimuli across culturally similar and dissimilar situations. Culture and Brain, in press.

Chen, Y., Shen, Y., Shi, Z., Zhang, X., Li, H., Xu, X., Guan, L., Han, S., Yang, J. (2020). Mortality salience impairs self-referential processing: Neurophysiological and behavioral evidence. Current Psychology, in press.

Ko, A., Pick, C. M., Kwon, J. Y., Barlev, M., Krems, J. A., Varnum, M. E., Neel, R., Peysha, M., Boonyasiriwat, W., Brandstätter, E., Vasquez, J.E.C., Galindo, O., David, D., de Felipe, R. P., Crispim, A. C., Fetvadijev, V.H., Fischer, R., Karl, J., Galdi, S., Gomez-Jacinto, L., Grossman, I., Gul, P., Hamamura, T., Han, S., Hitokoto, H., Hřebíčková, M., Graf, S., Johnson, J. L., Malanchuk, O., Murata, A., Na, J., Rizwan, M., Roth, R., Salgado, S. A., Sevincer, A. T., Stanciu, A., Suh, E. M., Talhelm, T., Uskul, A.K., Uz, I., Zambrano, D., Kenrick, D. T. (2020). Family matters: rethinking the psychology of human social motivation. Perspectives on Psychological Science, 15(1):173-201.

Qi, Y., Herrmann, M. J., Bell, L., Fackler, A., Han, S., Deckert, J., & Hein, G. (2020). The mere physical presence of another person reduces human autonomic responses to aversive sounds. Proceedings of the Royal Society B, 287(1919), 20192241.

Zhou, Y., Gao, T., Zhang, T., Li, W., Wu, T., Han, X., Han, S. (2020). Neural dynamics of racial categorization predicts racial bias in face recognition and altruism. Nature Human Behaviour, 4(1), 69-87.


Luo, S., Wu, B., Fan, X., Zhu, Y., Wu, X., & Han, S. (2019). Thoughts of death affect rewar learning by modulating salience network activity. NeuroImage, 202, 116068.

Pfabigan, D.M., Han, S. (2019). Converging electrophysiological evidence for a processing advantage of social over nonsocial feedback. Cognitive, Affective & Behavioral Neuroscience, 19(5):1170-1183.

Luo, S., Zhang, T., Li, W., Yu, M., Hein, G., Han, S. (2019). Interactions between oxytocin receptor gene and intergroup relationship on empathic neural responses to others’ pain. Social Cognitive and Affective Neuroscience, 14 (5), 505-517.

Li, W., Han, S. (2019). Behavioral and electctrophysiological evidence for enhanced sensitivity to subtle variations of pain expressions of same-race than other-race faces. Neuropsychologia, 129, 302-309.


Han, S. (2018). Neurocognitive basis of racial ingroup bias in empathy. Trends in Cognitive Sciences, 22, 400-421.

An. S., Han, X., Wu, B., Shi, Z., Marks, M., Wang, S., Wu, X., Han, S. (2018). Neural activation in response to the two sides of emotion. Neuroscience Letters, 684,140-144.

Feng, C., Yan, X., Huang, W., Han, S., Ma. Y. (2018). Neural representations of the multidimensional self in the cortical midline structures. NeuroImage, 183, 291-299.

Shi, Z., Han, S. (2018). Distinct effects of reminding mortality and physical pain on the default-mode activity and activity underlying self-reflection. Social Neuroscience, 13, 372-383.

Luo, S., Han, X., Du, D., Han, S. (2018). Physical coldness enhances racial in-group bias in empathy: Electrophysiological evidence. Neuropsychologia,116(Pt A), 117-125.

Qin, J., Shi, Z., Ma, Y., Han, S. (2018). Gender and neural substrates subserving implicit processing of death-related linguistic cues. Cognitive Process, 19, 63-71.

Liu, Y., Wu, B., Petti, C., Wu, X., Han, S. (2018). Self-construals moderate the associations between trait creativity and the social brain network. Neuropsychologia, 111, 284-291.

Fan, X., & Han, S. (2018). Distinct neural responses to one's own name under mortality threat in Christians and nonbelievers. Neuropsychologia, 108, 32-41.


Han, S. (2017). The social cultural brain: Cultural neuroscience approach to human nature. Oxford University Press, Oxford, UK.

Luo., S., Yu, D., Han, S. (2017). 5-HTTLPR moderates the association between interdependence and brain responses to mortality threats. Human Brain Mapping, 38, 6157-6171.

Mu, Y., Han, S., Gelfand, M. (2017). The role of gamma interbrain synchrony in social coordination when humans face territorial threats. Social Cognitive and Affective Neuroscience, 12, 1614-1623.

Han, X., He, K., Wu, B., Shi, Z., Liu, Y., Luo, L., Wei, K., Wu, X., Han. S. (2017). Empathy for pain motivates actions without altruistic effects: Evidence of motor dynamics and brain activity. Social Cognitive and Affective Neuroscience, 12, 893-901.

Han, X., Zhang, T., Wang, S., Han, S. (2017). Neural correlates of believing. NeuroImage, 156, 155-165.

Sheng, S., Du, N., Han, S. (2017). Degraded perceptual and affective processing of racial out-groups: An electrophysiological approach. Social Neuroscience, 12, 479-487.


Sheng, S., Du, N., Han, S. (2016). Degraded perceptual and affective processing of racial out-groups: An electrophysiological approach. Social Neuroscience, in press.

Mu, Y., Guo, C., Han, S. (2016). Oxytocin enhances inter-brain synchrony during social coordination in male adults. Social Cognitive and Affective Neuroscience, in press.

Han, S., Ma, Y., Wang, G. (2016). Shared neural representations of self and conjugal family members in Chinese brain. Culture and Brain, in press.

Ma, Y., Li, S., Wang, C., Liu, Y., Li, W., Yan, X., Chen, Q., Han, S. (2016). Distinct oxytocin effects on belief updating in response to desirable and undesirable feedback. Proceedings of the National Academy of Sciences, USA, 113, 9256-9261.

Han, X., Luo, S., Han, S. (2016). Embodied neural responses to others’ suffering. Cognitive Neuroscience, 7, 114-127.

Sheng, F., Han, X., Han, S. (2016). Dissociated neural representations of pain expressions of different races. Cerebral Cortex, 26, 1221-1233.

Yang, J., Xu, X., Chen, Y., Shi, Z., Han, S. (2016). Trait self-esteem and neural activities related to self-evaluation and social feedback. Scientific Reports, 6, 20274.

Luo, S., Yu, D., Han, S. (2016). Genetic and neural correlates of romantic relationship satisfaction. Social Cognitive and Affective Neuroscience, 11, 337-348.

Ma, Y., Wang, C., Luo, S., Li, B., Wager, T.D., Zhang, W., Rao, Y., Han, S. (2016). Serotonin transporter polymorphism alters citalopram effects on human pain responses to physical pain. NeuroImage, 135, 186-196.

Ma, Y., Shamay-Tsoory, S., Han, S., Zink, C. (2016). Oxytocin and social adaptation: Insights from neuroimaging studies of healthy and clinical populations. Trends in Cognitive Sciences, 20, 133-145.

Shi, Z., Ma, Y., Wu, B., Wu, X., Wang, Y., Han, S. (2016). Neural correlates of reflection on actual versus ideal self-discrepancy. NeuroImage, 124, 573-580.

Han, S., Humphreys, G. W. (2016). Self-construal: A cultural framework for brain function. Current Opinion in Psychology, 8, 10-14.

Mu, Y., Kitayama, S., Han, S., & Gelfand, M. J. (2016). Reply to Jacquet et al.: Culture and the neurobiology of norm violation detection. Proceedings of the National Academy of Sciences, USA, 113, E2874-2875.


Mu, Y., Kitayama, S., Han, S., Gelfand, M. (2015). How culture gets embrained: Cultural differences in event-related potentials of social norm violations. Proceedings of the National Academy of Sciences, USA, 112, 15348-15353.

Han. S., Ma. Y. (2015). Cultural neuroscience studies of self-reflection. In Joan Chiao, Shu-Chen Li, Rebecca Seligman, Robert Turner (eds.). The Oxford Handbook of Cultural Neuroscience. pp. 197-208. New York: Oxford University Press. .

Han, S., Ma. Y. (2015). A culture-behavior-brain loop model of human development.Trends in Cognitive Sciences, 9, 666-676.

Wang, C., Wu, B., Liu, Y., Wu, X., Han, S. Challenging emotional prejudice by changing self-concept: Priming independent self-construal reduces racial in-group bias in neural responses to other's pain. Social Cognitive and Affective Neuroscience, 10, 1195-1201.

Li, X., Liu, Y., Luo, S., Wu, B., Wu, X., Han, S. (2015). Mortality salience enhances racial in-group bias in empathic neural responses to others' suffering. NeuroImage, 116, 376-385.

Luo, S. Ma, Y., Liu, Y., Li, B., Wang, C., Shi, Z., Li, X., Zhang, W., Rao, Y., & Han, S. (2015). Interaction between oxytocin receptor polymorphism and interdependent culture on human empathy. Social Cognitive and Affective Neuroscience, 10, 1273-1281.

Luo, S., Li, B., Ma, Y., Zhang, W., Rao, Y., Han, S. (2015). Oxytocin receptor gene and racial ingroup bias in empathy-related brain activity. NeuroImage, 110, 22-31.

Ma, Y., Liu, Y., Rand, D., Heatherton, T., Han, S. (2015). Opposing oxytocin effects on inter-group cooperative behavior in intuitive and reflective minds. Neuropsychopharmacology, 40, 2379-2387.

Ma, Y., Li, B., Wang, C., Zhang, W., Rao., Y., Han, S. Genetic difference in acute citalopram effects on human emotional network. British Journal of Psychiatry, 206,385-392.

Mahmoodi, A., Bang, D., Olsen, K., Zhao, Y. A., Shi, Z., Broberg, K., Safavi, S., Han, S., Ahmadabadi, M. N., Frith, C, D., Roepstorff, A., Rees, G., Bahrami, B. (2015). An equality bias impairs collective decision-making across cultures. Proceedings of the National Academy of Sciences, USA.112,3835-3840.

Guan, L., Chen, Y., Xu, X., Qiao, L, Wei, L., Han, S., Yang, J., Liu, Y. (2015). Self-esteem buffers the mortality salience effect on the implicit self-face processing. Personality and Individual Differences 85, 77–85.

Meyer, M.L., Masten, C.L., Ma, L., Wang, C., Shi, Z., Eisenberger, N.I., Lieberman, M.D., Han, S. Differential neural activation to friends and strangers links interdependence to empathy. Culture and Brain, 3,321-338.

Han, S. (2015). Understanding cultural differences in human behavior: A cultural neuroscience approach. Current Opinion in Behavioral Sciences, 3, 68-72.

Freeman, J.B., Ma, Y., Barth, M., Young, S. G., Han, S., Ambady, N. (2015). The neural basis of contextual influences on face categorization. Cerebral Cortex, 25, 415-422.

Han S. (2015) Cultural Neuroscience. In: Arthur W. Toga, editor. Brain Mapping: An Encyclopedic Reference, vol. 3, pp. 217-220. Academic Press: Elsevier .


Sessa, P., Meconi, F., Han, S. (2014). Double dissociation of neural responses supporting perceptual and cognitive components of social cognition: Evidence from processing of others' pain. Scientific Reports, 4, doi:10.1038/srep07424.

Huang, S., Han, S. (2014). Shared beliefs enhance shared feelings: Religious/irreligious identifications modulate empathic neural responses. Social Neuroscience, 9, 639-649.

Zhou, B., Yang, S., Zhang, T., Mao, L., Han, S. (2014). Visual feature processing in the early visual cortex affects duration perception. Journal of Experimental Psychology: General. 143, 1893-1902.

Luo, S., Han, S. (2014). The association between an oxytocin receptor gene polymorphism and cultural orientations. Culture and Brain, 2, 89-107.

Korn, C. W., Fan, Y., Zhang, K., Wang, C., Han, S., Heekeren, H.R. (2014). Cultural influences on social feedback processing of character traits. Frontiers in Human Neuroscience, 8, 192.

Zaytseva, Y., Gutyrchik, E., Bao, Y., Han, S., Northoff, G., Welker, L., Meindl, T., Blautzik, J. (2014). Self processing in the brain: a paradigmatic fMRI case study with a professional singer. Brain and Cognition, 87C, 104-108.

Han, S., Ma,Y. (2014). Cultural differences in human brain activity: A quantitative meta-analysis. NeuroImage, 99, 293-300.

Zhang, X., Qiu, J., Zhang, Y., Han, S., Fang, F. (2014). Misbinding of color and motion in human visual cortex. Current Biology, 24, 1354-1360.

Luo, S., Shi, Z., Yang, X., Wang, X., Han, S. (2014). Reminders of mortality decrease midcingulate activity in response to others’ suffering. Social Cognitive and Affective Neuroscience, 9, 477-486.

Sheng, F., Liu, Q., Li, H., Fang, F., Han, S. (2014). Task modulations of racial bias in neural responses to others’ suffering. NeuroImage, 88, 263-270.

Jiang, C., Varnum, M.E.W., Hou, Y., Han, S. (2014). Distinct effects of self-construal priming on empathic neural responses in Chinese and Westerners. Social Neuroscience, 9, 130-138.

Wang, C., Ma, Y., Han, S. (2014). Self-construal priming modulates pain perception: Event-related potential evidence. Cognitive Neuroscience, 5, 3-9.

Varnum, M.E.W., Shi, Z., Chen, A., Qiu, J., Han, S. (2014). When “Your” reward is the same as “my” reward: Self-construal priming shifts neural responses to own vs. friends’ rewards. NeuroImage, 87, 164-169.

Ma, Y., Wang, C., Li, B., Zhang, W., Rao, Y., Han, S. (2014). Does self-construal predict activity in the social brain network? A genetic moderation effect. Social Cognitive and Affective Neuroscience, 9, 1360-1367.

Ma, Y., Li, B., Wang, C., Shi, Z., Sun, Y., Sheng, F., Zhang, Y., Zhang, W., Rao., Y., Han, S. (2014). 5-HTTLPR polymorphism modulates neural mechanisms of negative self-reflection. Cerebral Cortex, 24, 2421–2429.

Ma, Y., Bang, D., Wang, C., Allen, M., Frith, C., Roepstorff, A., Han, S. (2014). Sociocultural patterning of neural activity during self-reflection. Social Cognitive and Affective Neuroscience, 9, 73-80.


Liu, Y., Sheng, S., Woodcock, K. A., Han, S. Oxytocin effects on neural correlates of self-referential processing. Biological Psychology, 94, 380-387.

Woodcock, K. A., Yu, D., Liu, Y., Han, S. (2013). The presence of a culturally similar or dissimilar social partner affects neural responses to emotional stimuli. Socioaffective Neuroscience & Psychology, 3: 20500 -

Krämer, K., Bente, G., Luo, S., Pfeiffer, U. J., Han, S., Vogeley, K. (2013). Influence of ethnic group-membership and gaze direction on the perception of emotions: A cross-cultural study between Germany and China. PLoS ONE, 8(6):e66335. doi: 10.1371/journal.pone.0066335.

Liu, X., Shi, Z., Ma, Y., Qin, J., Han, S. (2013). Dynamic neural processing of linguistic cues related to death. PLoS ONE, 8(6): e67905. doi: 10.1371/journal.pone.0067905.

Shi, Z., Han, S. (2013). Transient and sustained neural responses to death-related linguistic cues. Social Cognitive and Affective Neuroscience, 8, 573-578.

Zuo, X., Han, S. (2013). Cultural experiences reduce racial bias in neural responses to others' suffering. Culture and Brain, 1, 34-46.

Han, S. (2013). Culture and Brain: a new journal. Culture and Brain, 1, 1-2.

Meyer, M. L., Masten, C. L., Ma, Y., Wang, C., Shi, Z., Eisenberger, N. I., Han, S. (2013). Empathy for the social suffering of friends and strangers recruits distinct patterns of brain activation. Social Cognitive and Affective Neuroscience, 8, 446-454.

Sui, J., Hong, Y., Liu, C. H. Humphreys, G. W., Han, S. (2013). Dynamic cultural modulation of neural responses to one’s own and friend’s faces. Social Cognitive and Affective Neuroscience, 8, 326-332.

Wang, C., Oyserman, D., Li, H., Liu, Q., Han, S. (2013). Accessible cultural mindset modulates default mode activity: Evidence for the culturally situated brain. Social Neuroscience, 8, 203-126.

Li, N., Ma, N., Liu, Y., He, X., Sun, D., Fu, X., Zhang, X., Han, S., Zhang, D. (2013). Resting state functional connectivity predicts impulsivity in economic decision-making. Journal of Neuroscience, 33, 4886-4895.

Han, S. (2013). How to identify mechanisms of cultural influences on human brain functions. Psychological Inquiry, 24, 37-41.

Kitayama, S., Chua, H. F., Tompson, S., Han, S. (2013). Neural mechanisms of dissonance: An fMRI investigation of choice justification. NeuroImage, 69, 206-212.

Han, S., Northoff, G., Vogeley, K., Wexler, B. E., Kitayama, S., Varnum, M. E. W. (2013). A cultural neuroscience approach to the biosocial nature of the human brain. Annual Review of Psychology, 64, 335–359.

Sheng, F., Liu, Y., Zhou, B., Zhou, W., Han, S. (2013). Oxytocin modulates the racial bias in neural responses to others’ suffering. Biological Psychology, 92, 380-386.

Mu, Y., Han, S. (2013). Neural oscillations dissociate between self-related attentional orienting versus evaluation. NeuroImage, 67, 247-256.

Freeman J. B., Ma, Y., Han, S., Ambady, N. (2013). Influences of culture and visual context on real-time social categorization. Journal of Experimental Social Psychology, 49, 206-210.

Han. S. (2013). Understanding the self: A cultural neuroscience perspective. In P. S Barnow and N. Balkir (eds.). Cultural Variation in Psychopathology (Chapter 2). pp.27-42, Cambridge, MA: Hogrefe Publishers.

Han, S. (2013). Cross-cultural variation in social cognition and the social brain. In D. L. Roberts & D. L. Penn (Eds.), Social cognition in schizophrenia: From evidence to treatment (Chapter 3). pp. 69-92, New York: Oxford University Press.


Ma, Y., Han, S. Functional dissociation of the left and right fusiform gyrus in self-face recognition. Human Brain Mapping, 33,2255-2267.

Ma, Y., Han, S. (2012). Is the self always better than a friend? Self-face recognition in Christians and atheists. PLoS ONE, 7(5): e37824. doi:10.1371/journal.pone. 0037824.

Sheng, F., Han, S. (2012). Manipulations of cognitive strategies and intergroup relationships reduce the racial bias in empathic neural responses. NeuroImage, 61, 786-797.

Lu, S., Cai, Y., Shen, M., Zhou, Y., Han, S. (2012). Alerting and orienting of attention without visual awareness. Consciousness and Cognition. 21, 928-938.

Wang, G., Mao, L., Ma, Y., Yang, X., Cao, J., Liu, X., Wang, J., Wang, X., Han, S. (2012). Neural representations of close others in collectivistic brains. Social Cognitive and Affective Neuroscience, 7, 222-229.

de Greck, M., Wang, G., Yang, X., Wang, X., Northoff, G., Han, S. (2012). Neural substrates underlying intentional empathy. Social Cognitive and Affective Neuroscience, 7, 135-144.

de Greck, M., Shi, Z., Wang, G., Zuo, X., Yang, X., Wang, X., Northoff, G., Han, S. (2012). Culture modulates brain activity during empathy with anger. NeuroImage, 59, 2871-2882.


Wang, J., Kitayama, S., Han, S. Sex difference in the processing of task-relevant and task-irrelevant social information: An event-related potential study of familiar face recognition. Brain Research, 1408, 41-51.

Liew, S., Han, S., Aziz-Zadeh, L. Familiarity modulates mirror neuron and mentalizing regions during intention understanding. Human Brain Mapping, in press.

Liew, S. L., Ma, Y., Han, S., Aziz-Zadeh, L. (2011). Who’s Afraid of the Boss: Cultural Differences in Social Hierarchies Modulate Self-face Recognition in Chinese and Americans. PLoS ONE, 6, e16901. doi:10.1371/journal.pone.0016901.

Qin, J., Kimel, S., Kitayama, S., Wang, X., Yang, X., Han, S. (2011). How choice modifies preference: Neural correlates of choice justification. NeuroImage, 55, 240-246.

Han, S., Mao, L., Qin, J., Friederici, A. D., Ge, J. (2011). Functional roles and cultural modulations of the medial prefrontal and parietal activity associated with causal attribution. Neuropsychologia, 49, 83-91.

Ma, Y., Yang, S., Han, S. Attitudes influence implicit racial face categorization in a perceptual task. Group Processes & Intergroup Relations, 14, 887-899.

Ma, Y., Wang, C., Han, S. (2011). Neural responses to perceived pain in others predict real-life monetary donations in different socioeconomic contexts. NeuroImage, 57, 1273-1280.

Ma, Y., Han, S. (2011). Neural representation of self-concept in sighted and congenitally blind adults. Brain, 134, 235-246.

Han, S., Ma., Y., & Sui, J. (2011). Self identity in sociocultural contexts: Implications from studies of self-face recognition. In Han, S., & Pöppel, E. (Eds.). Culture and Neural Frames of Cognition and Communication (On Thinking). pp 65-76, Berlin: Springer.

Zhang, L., Zhu, Y., & Han, S. (2011). The relation between the self and others: A transcultural neuroimaging approach. In Han, S., & Pöppel, E. (Eds.). Culture and Neural Frames of Cognition and Communication (On Thinking). pp 77-92, Berlin: Springer.


Han, S., Qin, J., Ma, Y. (2010). Neurocognitive processes of linguistic cues related to death. Neuropsychologia, 48, 3436-3442.

Mu, Y., Han, S. (2010). Neural oscillations involved in self-referential processing. NeuroImage, 53, 757-768.

Zhou, B., Qin, J., Mao, L., Pöppel, E., Han, S. (2010). Modulations of temporal perception by consciously and unconsciously perceived stimuli. Perception, 39, 900 – 908.

Lee, T.M.C., Ip, A.K.Y., Wang, K., Xi, H., Hu, P., Hu, C., Mak, H.K.F., Han, S., Chan, C.C.H., (2010). Faux pas deficits in people with medial frontal lesions as related to impaired understanding of a speaker's mental state. Neuropsychologia, 48, 1670-1676.

Han, S., Gu, X., Mao, L., Ge, J., Wang, G., Ma, Y. (2010). Neural substrates of self-referential processing in Chinese Buddhists. Social Cognitive and Affective Neuroscience, 5, 332-339.

Ma, Y., Han, S. (2010). Why respond faster to the self than others? An implicit positive association theory of self advantage during implicit face recognition. Journal of Experimental Psychology: Human Perception and Performance, 36, 619-633.

Ng, S. H., Han, S., Mao, L., Lai, J. C. L. (2010). Dynamic bicultural brains: A fMRI study of their flexible neural representation of self and significant others in response to culture priming. Asian Journal of Social Psychology, 13, 83-91.

Sun, Y., Gao, X., Han, S. (2010). Sex differences in face gender recognition: An event-related potential study. Brain Research, 1327:69-76.

Li. W., Han, S. (2010). Perspective taking modulates event related potentials to perceived pain. Neuroscience Letters, 469, 328-332.


Ge, J., Gu, X., Ji, M., Han, S. (2009). Neurocognitive processes of the religious leader in Christians. Human Brain Mapping, 30, 4012-4024.

Han, S., Fan, Y., Xu, X., Qin, J., Wu, B., Wang, X., Aglioti, S. M., Mao, L. (2009).Empathic neural responses to others’ pain are modulated by emotional contexts. Human Brain Mapping, 30, 3227-3237.

Sui, J., Liu, C. H., Han, S. (2009). Cultural difference in neural mechanisms of self-recognition. Social Neuroscience, 4, 402-411.

Han, S., Northoff, G. (2009). Understanding the self: a cultural neuroscience approach. Progress in Brain Research, 178, 203-212.

Xu, X., Zuo, X., Wang, X., Han, S. (2009). Do you feel my pain? Racial group membership modulates empathic neural responses, Journal of Neuroscience, 29, 8525-8529.

Mak, A.K.Y., Wong, M.M.C., Han, S., Lee, T.M.C. (2009). Gray matter reduction associated with emotion regulation in female outpatients with major depressive disorder: A voxel-based morphometry studyProgress in Neuro-Psychopharmacology & Biological Psychiatry, 33, 1184-1190.

Lu, S., Han, S. (2009). Attentional capture is contingent on the interaction between task demand and stimulus salience. Attention, Perception, & Psychophysics, 1, 1015-1026.

Ma, Y., Han, S. (2009). Self-face advantage is modulated by social threat — Boss effect on self-face recognition. Journal of Experimental Social Psychology, 45, 1048-1051.

Ma, Y., Ge, J., Xu, X., Fan, Y., Yang, S., Han. S. (2009). Asymmetric neurocognitive representation of ethnic in-group/out-group faces. Chinese Science Bulletin, 54, 2076-2081.

Sui, J., Liu, C. H., Wang, L., Han, S. (2009). Attentional orientation induced by temporarily established self-referential cues. Quarterly Journal of Experimental Psychology, Section A, 62, 844-849.

Casco, C., Campana, G., Han, S. & Guzzon, D. (2009). Psychophysical and electrophysiological evidence of independent facilitation by collinearity and similarity in texture grouping and segmentation. Vision Research, 49(6), 583-593.

Lin, Z., Han, S. (2009). Self-construal priming modulates the scope of visual attention. Quarterly Journal of Experimental Psychology, Section A, 62, 802-813.

Qin, J., Lee, T. M. C., Han, S. (2009). Theta and alpha oscillations linked to risk Identifications. Brain Research, 1269, 125-134.

Qin, J., Lee, T. M. C., Wang, F., Mao, L., Han, S. (2009). Neural activities underlying environmental and personal risk identification tasks. Neuroscience Letters, 455, 110-115.

Qin, J., Han, S. (2009). Parsing neural mechanisms of social and physical risk identifications. Human Brain Mapping, 30, 1338-1351.

Qin, J., Han, S. (2009). Neurocognitive mechanisms underlying identification of environmental risks. Neuropsychologia, 47, 397-405.

Han, S. (2009). Religious belief and neurocognitive processes of the self. E.Voland & W. Schiefenhövel (eds). The biological evolution of religious mind and behaviour, The Frontiers Collection, pp.195-204, Springer-Verlag Berlin Heidelberg.

Ng. S. H., Han, S. (2009). The bilcultural self and the bilcultural brain. In Wyer, R. S., Chiu, C.-y., & Hong, Y.-y (Eds.) Problems and solutions in cross-cultural theory, research and application. pp 329-342, New York: Psychology Press.


Han, S., Northoff, G. (2008). Reading direction and culture. Nature Review Neuroscience, 9, 965.

He, X., Humphreys, G. W., Fan, S., Chen, L., Han, S. (2008). Differentiating spatial and object-based effects on attention: An event-related brain potential study with peripheral cueing. Brain Research, 1245, 116-125.

Han, S., Fang, F. (2008). Linking neural activity to mental processes. Brain Imaging and Behavior, 2, 242-248.

Mu, Y., Fan, Y., Mao, L., Han, S. (2008). Event-related theta and alpha oscillations mediate empathy for pain. Brain Research. 1234, 128-136.

Zhu, Y., Han, S. (2008). Cultural differences in the self: From philosophy to psychology and neuroscience. Social and Personality Psychology Compass. 2, 1799-1811.

Ge. J., Han, S. (2008). Distinct neurocognitive strategies for comprehensions of human and artificial intelligence. PLoS ONE 3(7): e2797.doi:10.1371/journal. pone.0002797.

Han, S., Northoff, G. (2008). Culture-sensitive neural substrates of human cognition: A transcultural neuroimaging approach. Nature Review Neuroscience, 9, 646-654.

Lee, T. M. C., Chan, C. C. H., Han, S., Leung, A. W. S., Fox, P. T., & Gao, J.-H. (2008). An event-related fMRI study on risk taking by healthy individuals of high or low impulsiveness. Neuroscience Letters, 438, 138-141

Han, S., Gao, X., Humphreys, G. W., Ge, J. (2008). Neural processing of threat cues in social environments. Human Brain Mapping, 29, 945-957.

Lin, Z., Lin., Y., Han, S. (2008). Self-construal priming modulates visual activity underlying global/local perception. Biological Psychology. 77, 93-97.

Han, S., Mao, L., Gu, X., Zhu, Y., Ge, J., Ma, Y. (2008). Neural consequences of religious belief on self-referential processing. Social Neuroscience, 3, 1-15.

Fan, Y., Han, S. (2008). Temporal dynamic of neural mechanisms involved in empathy for pain: An event-related brain potential study. Neuropsychologia, 46, 160-173.

Han, S., Fan, Y., Mao, L. (2008). Gender difference in empathy for pain: An electrophysiological investigation. Brain Research, 1192, 85-93.


Zhu, Y., Zhang, Li., Fan, J., Han. S. (2007). Neural basis of cultural influence on self representation. Neuroimage, 34, 1310-1317.

Mao, L., Zhou, B., Zhou, W., Han, S. (2007). Neural correlates of covert orienting of visual spatial attention along vertical and horizontal dimensions. Brain Research. 1136, 142-153.

Sui, J., Han, S. (2007). Self-construal priming modulates neural substrates of self-awareness. Psychological Science. 18, 861-866.

Gu, X., Han, S. (2007). Attention and reality constraints on the neural processes of empathy for pain. Neuroimage. 36, 256-267.

Jiang, Y., Han, S. (2007). Perceptual gain and perceptual loss: Distinct neural mechanisms of audiovisual interactions. International Journal of Magnetic Resonance Imaging. 1(1), 3-14.

Wu, X., Chen, X., Li, Z., Han, S., Zhang, D. (2007). Binding of verbal and spatial information in human working memory involves large scale neural syncronization at theta frequency. Neuroimage, 35, 1654-1662.

Zhang, X., Han, S. (2007). Global perception depends on coherent work of bilateral visual cortices: TMS studies. Science in China, series C. 50(4), 557-565.

Gu, X., Han, S. (2007). Neural substrates underlying evaluation of pain in actions depicted in words. Behavioural Brain Research, 181, 218-223.

Han, S., Humphreys, G. W. (2007). The fronto-parietal network and top-down modulation of perceptual grouping. Neurocase, 13, 278-289.

Qin, J., Han, S. (2007). The role of parietal cortex in global/local processing of hierarchical stimuli: A TMS study. NeuroReport, 18, 1921-1924.

Han, S., Jiang, Y., Humphreys, G. W. (2007). Watching cartoons activates the medial prefrontal cortex in children. Chinese Science Bulletin, 52, 3371-3375.


Mao, L., Zhang, X., Chen, J., Gu, H., Han, S. (2006). Neural substrates of global perception are modulated by local element grouping. Chinese Science Bulletin, 51 (6), 1-6.

Han, S., Jiang, Y., Mao, L. (2006). Right hemisphere dominance in perceiving coherence of visual events. Neuroscience Letters, 398 (1-2): 18-21.

Sui, J., Zhu, Y., Han, S. (2006). Self-face recognition in attended and unattended conditions: an ERP study. NeuroReport, 17(4):423-427.

Han, S., Jiang, Y. (2006). Neural correlates of within-level and across-level attention to multiple compound stimuli. Brain Research, 1076: 193-197.

Li, Z., Bao, M., Chen, X., Zhang, D., Han, S., He, S., Hu, X. (2006). Attention shift in human verbal working memory: priming contribution and dynamic brain activation. Brain Research, 1078: 131-142.

Han, S., Glyn, W. Humphreys (2005). Perceptual organization at attended and unattended locations. Science in China, series C, 48(2), 106-116.

Han, S., Jiang, Y., Humphreys, G. W., Zhou, T., & Cai, P. (2005). Distinct neural substrates for the perception of real and virtual visual worlds. Neuroimage, 24, 928-935.

Han, S., Wan, X., & Humphreys, G. W. (2005). Shifts of spatial attention in a perceived 3-D space. Quarterly Journal of Experimental Psychology, Section A, 58A, 753-764.

Han, S., Jiang, Y., Mao L., Humphreys, G. W., Gu, H. (2005). Attentional modulation of perceptual grouping in human visual cortex: fMRI studies. Human Brain Mapping, 25, 424-432.

Jiang, Y., Han, S. (2005). Neural mechanisms of global/local processing of bilateral visual inputs: an ERP study. Clinical Neurophysiology, 116, 1444-1454.

Wu, Y., Chen, J., Han, S. (2005). Neural mechanisms of attentional modulation of perceptual grouping by collinearity. NeuroReport, 16, 567-570.

Han, S., Jiang, Y., Mao L., Humphreys, G. W., Qin, J. (2005). Attentional modulation of perceptual grouping in human visual cortex: ERP studies. Human Brain Mapping, 26, 199-209.

2003- 2004

Han, S., Jiang, Y., Gu, H., Rao, H., Mao, L., Cui, Y., Zhai, R. (2004). The role of human parietal cortex in attention networks. Brain, 127: 650-659.

Mao, L., Han, S., Guo, C., & Jiang, Y. (2004). Neural mechanisms of perceptual grouping in human visual cortex. Chinese Science Bulletin, 49(8): 819-823.

Han, S., Jiang, Y., & Gu, H. (2004). Neural substrates differentiating global/local processing of bilateral visual inputs. Human Brain Mapping, 22 (4): 321-328.

Liu, L., Wang, K., Liao, B., Xu, L., & Han, S. (2004) Perceptual salience of global structures and the crowding effect in amblyopia. Clinical and Experimental Ophthalmology, 242, 566-570.

Rao, H., Han, S., Jiang, Y., Xue, Y., Gu, H., Cui, Y., Gao, D. (2004). Engagement of the prefrontal cortex in representational momentum: An fMRI study. NeuroImage, 23 (1), 98-103.

Han, S. (2004). Interactions between proximity and similarity grouping: An ERP study. Neuroscience Letters, 367 (1), 40-43.

Han, S., Jiang, Y. (2004). The parietal cortex and attentional modulations of activities of the visual cortex. NeuroReport, 15, 2275-2280.

Han, S., Wang, C., Zhou, L. (2004). Global and local processing under attended and unattended conditions. Acta Psychologica Sinica, 36(4), 410-416.

Zhou, B., Zhang, J. X., Tan, L. H., & Han, S (2004). Spatial congruence in working memory: an ERP study. NeuroReport, 15, 2795-2799.

Han, S., & Humphreys, G.W. (2003). Relationship between uniform connectedness and proximity in perceptual grouping. Science in China, series C, 46(2), 113- 126.

Han, S., & He, X. (2003). Modulation of neural activities by enhanced local selection in the processing of compound stimuli. Human Brain Mapping, 19(4), 273-281.

Han, S., Yund, E. W., & Woods, D. L. (2003). An ERP study of the global precedence effect: the role of spatial frequency. Clinical Neurophysiology, 114 (10), 1850-1865.

Han, S., Weaver, J., Murray, S., Yund, E. W., & Woods, D. L. (2003). Modulation of human global/local perception by low spatial frequency filtering. Chinese Sceince Bulletin. 48 (21), 2336- 2339.

Jiang, Y., Han, S. (2003). Perceptual grouping by proximity and similarity in the peripheral vision. Acta Psychologica Sinica, 35 (supplement): 70-78.

2001- 2002

Han, S., & Humphreys, G.W. (2002). Segmentation and selection contribute to local processing in hierarchical analysis. Quarterly Journal of Experimental Psychology, Section A, 55(1), 5-21.

Han, S., Ding, Y., & Song, Y. (2002). Neural mechanisms of perceptual grouping in humans as revealed by high density event related potentials. Neuroscience Letters, 319, 29-32.

Han, S., Weaver, J., Murray, S., Yund, E. W., & Woods, D. L. (2002). Hemispheric asymmetry in global/local processing: Effects of stimulus position and spatial frequency. NeuroImage, 17, 1290 -1299.

Chen, H., Li, S., Han, S., Wang, X. (2002). The research on visual global and local cognitive function of patients with Parkinson’s disease. Chinese Journal of Clinical Rehabilitation. 6(11): 1694- 1695.

Han, S., He, X., Yund, E. W., & Woods, D. L. (2001). Attentional selection in the processing of hierarchical patterns: An ERP study. Biological Psychology, 56 (2-3), 113-130.

Han, S., Song, Y., Ding, Y., Yund, E. W., & Woods, D. L. (2001). Neural substrates for visual perceptual grouping in humans. Psychophysiology, 38(6), 926-935.


Han, S., Liu, W., Yund, E. W., & Woods, D. L. (2000). Interactions between spatial attention and global/local feature selection: An ERP study. NeuroReport. 11(12), 2753-2758.

Han, S., He, X., & Woods, D. L. (2000). Hierarchical processing and level-repetition effect as indexed by early brain potentials. Psychophysiology, 37(6), 817-830.

Han, S., Fan, S., Chen, L., & Zhuo, Y.(1999). Modulation of brain activities by hierarchical processing: A high-density ERP study. Brain Topography, 11(3):171-183.

Han, S., Humphreys, G.W., & Chen, L. (1999). Uniform connectedness and classical Gestalt principles of perceptual grouping. Perception & Psychophysics, 61(4):661-674.

Han, S., Humphreys, G.W., & Chen, L. (1999). Parallel and competitive processes in hierarchical analysis: Perceptual grouping and encoding of closure. Journal of Experimental Psychology: Human Perception and Performance, 25(5), 1411-1432.

Han, S., & Humphreys, G.W. (1999). Interactions between perceptual organization based on Gestalt laws and those based on hierarchical processing. Perception & Psychophysics, 61(7), 1287-1298.


Han S, Fan. S, Chen, L., & Zhuo, Y. (1997). On the different processing of wholes and parts: A psychophysiological study. Journal of Cognitive Neuroscience, 9(5): 686-697.

Han, S., Humphreys, G.W., & Wang, R. (1997). Representation of the centre of a perceptual group in neglect: A case study. Neurocase, 3, 365-374.

1993- 1996

Han S, Chen L. (1996). Processing of global and local properties — An analysis with event- related brain potentials. Science in China, series C, 39(2), 179-188.

Han, S., Lu, J., Fan, S., Chen, L., & Zhuo, Y.(1996). The role of similarity in processing of compound patterns: An ERP study. Chinese Science Bulletin, 41(6):512-515.

Han, S. & Chen, L (1994). Different attentional effects on detection of orientation and color. Chinese Science Bulletin. 39(12). 1045-1048.

Han, S. & Chen, L. (1993). The different roles of attention oriented with peripheral cues from that of attention oriented with central cues: a topological explanation. In Wang Su (Ed.), Proceedings of the Second Afro-Asian Psychological Congress. Beijing University Press, p55-60.

Han, S. & Chen, L. (1993). The effects of central and peripheral cues on visual discriminability: A topological explanation. In Valenti, S.S. & Pittenger,J.B. (Eds) Studies in perception and action II: Posters presented at the VII International Conference on Event Perception and Action. Hillsdale, NJ: Lawrence Erlbum Associates, Inc, p61-65.

Han, S. (1993). Attention and the detection of color and orientation. In Valenti, S.S. & Pittenger,J.B. (Eds) Studies in perception and action II: Posters presented at the VII International Conference on Event Perception and Action. Hillsdale,NJ:Lawrence Erlbum Associates,Inc, p66- 68.